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* Key threats to chimpanzees are poaching and habitat destruction due to logging and shifting agriculture.
* Key threats to chimpanzees are poaching and habitat destruction due to logging and shifting agriculture.
* A REDD+ project is being developed and implemented at the site with the support of the Royal Society for the Protection of Birds, which aims to ensure funds for the continuation of conservation work in the park and improve the management of natural resources.
* A REDD+ project is being developed and implemented at the site with the support of the Royal Society for the Protection of Birds, which aims to ensure funds for the continuation of conservation work in the park and improve the management of natural resources.




= Site characteristics =
= Site characteristics =


The [https://www.golarainforest.org/the-forest site] consists of three forest blocks (Gola North, Central, and South) situated in southeast Sierra Leone, which are linked to Liberia’s Gola National Park, forming a [https://www.africa-eu-partnership.org/en/success-stories/trans-boundary-peace-park-sierra-leone-and-liberia transboundary peace park]. The forests in Gola Rainforest National Park are a catchment for the Moro, Mahoi, Mano and Moa rivers, providing an important water source for nearby villages (Barca et al. 2018). The area forms part of the Upper Guinean forest ecosystem, a biodiversity hotspot with a large number of endemic species. Close to 1,000 plant species have been identified in [https://www.golarainforest.org/the-forest Gola Rainforest National Park], half of which are endemic to the Upper Guinea forest ecosystem. In addition to the western chimpanzee (''Pan troglodytes verus''), other [https://www.golarainforest.org/wildlife threatened species] inhabit the area, including pygmy hippos (''Choeropsis liberiensis''), pangolins, red colobus (''Piliocolobus badius''), and Jentink’s duiker (''Cephalophus jentinki''). Gola Rainforest National Park is also classified as an Important Bird Area (BirdLife International 2019a) and has been included in the tentative list of [https://whc.unesco.org/en/tentativelists/5747/ World Heritage Sites].
The [https://www.golarainforest.org/the-forest site] consists of three forest blocks (Gola North, Central, and South) situated in southeast Sierra Leone, which are linked to Liberia’s Gola National Park, forming a [https://www.africa-eu-partnership.org/en/success-stories/trans-boundary-peace-park-sierra-leone-and-liberia transboundary peace park]. The forests in Gola Rainforest National Park are a catchment for the Moro, Mahoi, Mano and Moa rivers, providing an important water source for nearby villages ([[#R00634|Barca et al. 2018]]). The area forms part of the Upper Guinean forest ecosystem, a biodiversity hotspot with a large number of endemic species. Close to 1,000 plant species have been identified in [https://www.golarainforest.org/the-forest Gola Rainforest National Park], half of which are endemic to the Upper Guinea forest ecosystem. In addition to the western chimpanzee (''Pan troglodytes verus''), other [https://www.golarainforest.org/wildlife threatened species] inhabit the area, including pygmy hippos (''Choeropsis liberiensis''), pangolins, red colobus (''Piliocolobus badius''), and Jentink’s duiker (''Cephalophus jentinki''). Gola Rainforest National Park is also classified as an Important Bird Area ([[#R00636|BirdLife International 2019a]]) and has been included in the tentative list of [https://whc.unesco.org/en/tentativelists/5747/ World Heritage Sites].


'''Table 1. Basic site information for Gola Rainforest National Park'''
'''Table 1. Basic site information for Gola Rainforest National Park'''
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= Ape status =
= Ape status =


The chimpanzee population in Gola Rainforest National Park is estimated to be 270 individuals, which is equivalent to almost 5% of Sierra Leone’s estimated population (Brncic et al. 2010). The population trend is unknown.
The chimpanzee population in Gola Rainforest National Park is estimated to be 270 individuals, which is equivalent to almost 5% of Sierra Leone’s estimated population ([[#R00638|Brncic et al. 2010]]). The population trend is unknown.


'''Table 2. Ape population estimates reported for Gola Rainforest National Park'''
'''Table 2. Ape population estimates reported for Gola Rainforest National Park'''
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|-
|-
|''Pan troglodytes verus''
|''Pan troglodytes verus''
|2006-2007
|2015-2016
|
|
|0.102
|0.6
|
|
|
|
|Gola West, East, North, and an extension of Gola North
|Gola Rainforest National Park
|Line transects
|Line transects
|
|
|Klop et al. 2008
|[[#R00634|Barca et al. 2018]]
|Survey effort: 245.28 km. No evidence of chimpanzees found in Gola West
|96 chimpanzee nests were recorded
|
|SLE0012
|-
|-
|''Pan troglodytes verus''
|''Pan troglodytes verus''
|2009
|2009
|
|
|
|0.76
|0.27 (0.18-0.42)
|0.25 (0.14-0.43)
|305 (203-458)
|270 (159-468)
|Gola forest reserves and potential corridors
|Gola forest reserves and potential corridors
|Line transects
|Line transects
|
|
|Ganas 2009 as cited in Brncic et al. 2010
|[[#R00638|Brncic et al. 2010]]
|Survey effort: 173 km
|Population estimate based on a re-analysis of data presented in Ganas 2009
|
|SLE0002, SLE0003<div class="mw-collapsible mw-collapsed" data-expandtext="+13 more" data-collapsetext="less">SLE0004, SLE0014, SLE0015, SLE0017, SLE0018, SLE0021, SLE0024, SLE0025, SLE0026, SLE0029, SLE0032, SLE0036, SLE0037</div>
|-
|-
|''Pan troglodytes verus''
|''Pan troglodytes verus''
|2009
|2009
|
|
|0.76
|
|0.25 (0.14-0.43)
|0.27 (0.18-0.42)
|270 (159-468)
|305 (203-458)
|Gola forest reserves and potential corridors
|Gola forest reserves and potential corridors
|Line transects
|Line transects
|
|
|Brncic et al. 2010
|Ganas 2009 as cited in [[#R00638|Brncic et al. 2010]]
|Population estimate based on a re-analysis of data presented in Ganas 2009
|Survey effort: 173 km
|
|SLE0002, SLE0003<div class="mw-collapsible mw-collapsed" data-expandtext="+13 more" data-collapsetext="less">SLE0004, SLE0014, SLE0015, SLE0017, SLE0018, SLE0021, SLE0024, SLE0025, SLE0026, SLE0029, SLE0032, SLE0036, SLE0037</div>
|-
|-
|''Pan troglodytes verus''
|''Pan troglodytes verus''
|2015-2016
|2006-2007
|
|
|0.6
|0.102
|
|
|
|
|Gola Rainforest National Park
|Gola West, East, North, and an extension of Gola North
|Line transects
|Line transects
|
|
|Barca et al. 2018
|[[#R00642|Klop et al. 2008]]
|96 chimpanzee nests were recorded
|Survey effort: 245.28 km. No evidence of chimpanzees found in Gola West
|
|
|}
|}
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= Threats =
= Threats =


Approximately 28% of Gola Forest has been subject to logging (Klop et al. 2008). Large-scale logging operations in Gola started in the 1960s and continued until the late 1980s (Klop et al. 2008). Forestry Industries Corporation (FIC) and the Sierra Leone Timber Industry and Plantation Company (Sileti) were the two main companies responsible for the exploitation of timber in Gola (Klop et al. 2008). The resulting impacts from these logging operations include an extensive road network in Gola North, and a large clearing in Gola East. During the 1991-2001 civil war in Sierra Leone, the Revolutionary United Front set up camps near Gola forest; however, a survey that compared wildlife abundances before and after the war found no evidence of serious negative impacts from the war (Lindsell et al. 2011). [https://www.birdlife.org/worldwide/projects/forests-hope-site-gola-rainforest-national-park-sierra-leone Current threats] in the area include small-scale logging, bushmeat hunting, and shifting agriculture.
Approximately 28% of Gola Forest has been subject to logging ([[#R00642|Klop et al. 2008]]). Large-scale logging operations in Gola started in the 1960s and continued until the late 1980s ([[#R00642|Klop et al. 2008]]). Forestry Industries Corporation (FIC) and the Sierra Leone Timber Industry and Plantation Company (Sileti) were the two main companies responsible for the exploitation of timber in Gola ([[#R00642|Klop et al. 2008]]). The resulting impacts from these logging operations include an extensive road network in Gola North, and a large clearing in Gola East. During the 1991-2001 civil war in Sierra Leone, the Revolutionary United Front set up camps near Gola forest; however, a survey that compared wildlife abundances before and after the war found no evidence of serious negative impacts from the war ([[#R00643|Lindsell et al. 2011]]). [https://www.birdlife.org/worldwide/projects/forests-hope-site-gola-rainforest-national-park-sierra-leone Current threats] in the area include small-scale logging, bushmeat hunting, and shifting agriculture.


'''Table 3. Threats to apes reported for Gola Rainforest National Park'''
'''Table 3. Threats to apes reported for Gola Rainforest National Park'''
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!Year of threat
!Year of threat
|-
|-
|5 Biological resource use
|2 Agriculture & aquaculture
|5.3 Logging & wood harvesting
|2.1 Annual & perennial non-timber crops
|High (more than 70% of population affected)
|Present (unknown severity)
|Large-scale logging operations between 1960s and 1980s (Klop et al. 2008), Small-scale logging by local communities (BirdLife International 2019a)
|Shifting agriculture due to increasing population growth around the area and scarcity of fertile land ([[#R00637|BirdLife International 2019b]])
|1960-1989
|Ongoing (2019)
|-
|-
|4 Transportation & service corridors
|4 Transportation & service corridors
|4.1 Roads & railroads
|4.1 Roads & railroads
|High (more than 70% of population affected)
|High (more than 70% of population affected)
|Extensive road network in Gola North as a result of large-scale logging activities (Klop et al. 2008), which facilitates access to the forest.
|Extensive road network in Gola North as a result of large-scale logging activities ([[#R00642|Klop et al. 2008]]), which facilitates access to the forest.
|Ongoing (2008)
|Ongoing (2008)
|-
|-
|2 Agriculture & aquaculture
|5 Biological resource use
|2.1 Annual & perennial non-timber crops
|5.3 Logging & wood harvesting
|Present (unknown severity)
|High (more than 70% of population affected)
|Shifting agriculture due to increasing population growth around the area and scarcity of fertile land (BirdLife International 2019b)
|Large-scale logging operations between 1960s and 1980s ([[#R00642|Klop et al. 2008]]), Small-scale logging by local communities ([[#R00636|BirdLife International 2019a]])
|Ongoing (2019)
|1960-1989
|}
|}


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= Conservation activities =
= Conservation activities =


A conservation program began in 1990 by the Forestry Division of the government of Sierra Leone and the Conservation Society for Sierra Leone (CSSL). Since then, the Royal Society for the Protection of Birds (RSPB) became a supporter of the conservation projects in Gola, and the site is part of RSPB’s [http://www.birdlife.org/worldwide/programmes/forests-hope Forests of Hope Programme]. Conservation activities were interrupted by the civil war but resumed after the return of peace in 2002 (Lindsell et al. 2011).
A conservation program began in 1990 by the Forestry Division of the government of Sierra Leone and the Conservation Society for Sierra Leone (CSSL). Since then, the Royal Society for the Protection of Birds (RSPB) became a supporter of the conservation projects in Gola, and the site is part of RSPB’s [http://www.birdlife.org/worldwide/programmes/forests-hope Forests of Hope Programme]. Conservation activities were interrupted by the civil war but resumed after the return of peace in 2002 ([[#R00643|Lindsell et al. 2011]]).


A [https://www.rspb.org.uk/our-work/conservation/projects/gola-rainforest-sierra-leones-first-rainforest-national-park/ REDD+ project] is currently under development at the site. The project will use funds obtained from the sale of carbon credits to improve the management of the park, fund long-term conservation work, develop sustainable farming practices, and the establishment of a research station in the park.
A [https://www.rspb.org.uk/our-work/conservation/projects/gola-rainforest-sierra-leones-first-rainforest-national-park/ REDD+ project] is currently under development at the site. The project will use funds obtained from the sale of carbon credits to improve the management of the park, fund long-term conservation work, develop sustainable farming practices, and the establishment of a research station in the park.
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!Year of activity
!Year of activity
|-
|-
|5 Protection & restoration
|1 Development impact mitigation
|5.2 Legally protect ape habitat
|1.4 Farm more intensively and effectively in selected areas and spare more natural land
|The site was designated as National Park in 2010 (Gola Rainforest National Park 2019b)
|Develop sustainable farming practices ([[#R00637|BirdLife International 2019b]])
|
|
|2010-Ongoing (2019)
|Ongoing (2019)
|-
|-
|1 Development impact mitigation
|1 Development impact mitigation
|1.8 Use set-aside areas of natural habitat (e.g., mining, logging, agricultural areas)
|1.8 Use set-aside areas of natural habitat (e.g., mining, logging, agricultural areas)
|Bordering the site, cocoa trees are strategically planted alongside tree and plant species that increase soil fertility and attract birds (BirdLife International 2018)
|Bordering the site, cocoa trees are strategically planted alongside tree and plant species that increase soil fertility and attract birds ([[#R00635|BirdLife International 2018]])
|
|
|Ongoing (2018)
|Ongoing (2018)
|-
|1 Development impact mitigation
|1.4 Farm more intensively and effectively in selected areas and spare more natural land
|Develop sustainable farming practices (BirdLife International 2019b)
|
|Ongoing (2019)
|-
|-
|2 Counter-wildlife crime
|2 Counter-wildlife crime
|2.3 Conduct regular anti-poaching patrols
|2.3 Conduct regular anti-poaching patrols
|Forest guards patrol and monitor the forest (Gola Rainforest National Park 2019a)
|Forest guards patrol and monitor the forest ([[#R00640|Gola Rainforest National Park 2019a]])
|
|
|Ongoing (2019)
|Ongoing (2019)
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|4 Education & awareness
|4 Education & awareness
|4.1 Educate local communities about apes and sustainable use
|4.1 Educate local communities about apes and sustainable use
|The Gola Rainforest Conservation Center offers environmental education and conservation awareness to school classes and local communities (Gola Rainforest National Park 2019a)
|The Gola Rainforest Conservation Center offers environmental education and conservation awareness to school classes and local communities ([[#R00640|Gola Rainforest National Park 2019a]])
|
|
|Ongoing (2019)
|Ongoing (2019)
|-
|5 Protection & restoration
|5.2 Legally protect ape habitat
|The site was designated as National Park in 2010 ([[#R00641|Gola Rainforest National Park 2019b]])
|
|2010-Ongoing (2026)
|-
|-
|7 Economic & other incentives
|7 Economic & other incentives
|7.2 Provide non-monetary benefits to local communities for sustainably managing their forest and its wildlife (e.g., better education, infrastructure development)
|7.2 Provide non-monetary benefits to local communities for sustainably managing their forest and its wildlife (e.g., better education, infrastructure development)
|RSPB is developing a REDD+ project which aims to provide funds for the long-term management of the site and support local people in the sustainable management of resources (RSPB 2019)
|RSPB is developing a REDD+ project which aims to provide funds for the long-term management of the site and support local people in the sustainable management of resources ([[#R00644|RSPB 2019]])
|
|
|Ongoing (2019)
|Ongoing (2019)
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|2 Resources and capacity
|2 Resources and capacity
|2.5 Lack of equipment/transportation
|2.5 Lack of equipment/transportation
|Brncic et al. 2010
|[[#R00638|Brncic et al. 2010]]
|
|
|}
|}
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= Research activities =
= Research activities =


Several surveys have been done at the site to monitor wildlife and estimate species abundances (e.g., Brncic et al. 2010, Klop et al. 2008, Davies et al. 2008), and the nest ecology of chimpanzees has also been studied at the site (Barca et al. 2018).
Several surveys have been done at the site to monitor wildlife and estimate species abundances (e.g., [[#R00638|Brncic et al. 2010]], [[#R00642|Klop et al. 2008]], [[#R00639|Davies et al. 2008]]), and the nest ecology of chimpanzees has also been studied at the site ([[#R00634|Barca et al. 2018]]).




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= Exposure to climate change impacts =
= Exposure to climate change impacts =


As part of a study on the exposure of African great ape sites to climate change impacts, Kiribou et al. subm. extracted climate data and data on projected extreme climate impact events for the site. Climatological characteristics were derived from observation-based climate data provided by the Inter-Sectoral Impact Model Intercomparison Project (ISIMIP, www.isimip.org). Parameters were calculated as the average across each 30-year period. For 1981-2010, the EWEMBI dataset from ISIMIP2a was used. For the two future periods (2021-2050 and 2071-2099) ISIMIP2b climate data based on four CMIP5 global climate models were used. For future projections, two Representative Concentration Pathways (RCP) were used. RCP 2.6 is a scenario with strong mitigation measures in which global temperatures would likely rise below 2C. RCP 6.0 is a scenario with medium emissions in which global temperatures would likely rise up to 3C by 2100. For the number of days with heavy precipitation events, the 98th percentile of all precipitation days (>1mm/d) was calculated for the 1979-2013 reference period as a threshold for a heavy precipitation event. Then, for each year, the number of days above that threshold was derived. The figures on temperature and precipitation anomaly show the deviation from the mean temperature and mean precipitation for the 1979-2013 reference period.
As part of a study on the exposure of African great ape sites to climate change impacts, [[#R02204|Kiribou et al. (2024)]] extracted climate data and data on projected extreme climate impact events for the site. Climatological characteristics were derived from observation-based climate data provided by the Inter-Sectoral Impact Model Intercomparison Project (ISIMIP, www.isimip.org). Parameters were calculated as the average across each 30-year period. For 1981-2010, the EWEMBI dataset from ISIMIP2a was used. For the two future periods (2021-2050 and 2071-2099) ISIMIP2b climate data based on four CMIP5 global climate models were used. For future projections, two Representative Concentration Pathways (RCP) were used. RCP 2.6 is a scenario with strong mitigation measures in which global temperatures would likely rise below 2C. RCP 6.0 is a scenario with medium emissions in which global temperatures would likely rise up to 3C by 2100. For the number of days with heavy precipitation events, the 98th percentile of all precipitation days (>1mm/d) was calculated for the 1979-2013 reference period as a threshold for a heavy precipitation event. Then, for each year, the number of days above that threshold was derived. The figures on temperature and precipitation anomaly show the deviation from the mean temperature and mean precipitation for the 1979-2013 reference period.


The estimated exposure to future extreme climate impact events (crop failure, drought, river flood, wildfire, tropical cyclone, and heatwave) is based on a published dataset by Lange et al. 2020 derived from ISIMIP2b data. The same global climate models and RCPs as described above were used. Within each 30-year period, the number of years with an extreme event and the average proportion of the site affected were calculated (see Kiribou et al. subm. for details).
The estimated exposure to future extreme climate impact events (crop failure, drought, river flood, wildfire, tropical cyclone, and heatwave) is based on a published dataset by Lange et al. 2020 derived from ISIMIP2b data. The same global climate models and RCPs as described above were used. Within each 30-year period, the number of years with an extreme event and the average proportion of the site affected were calculated ([[#R02204|Kiribou et al. 2024]]).


'''Table 8. Estimated past and projected climatological characteristics in Gola Rainforest National Park'''
'''Table 8. Estimated past and projected climatological characteristics in Gola Rainforest National Park'''
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= References =
= References =


Barca, B., Turay, B.S., Kanneh, B.A., Tayleur, C. 2018. Nest ecology and conservation of Western Chimpanzees (Pan troglodytes verus) in Gola Rainforest National Park, Sierra Leone. Primate Conservation (32): 133-139.
<span id="R00634"></span>Barca, B., Turay, B.S., Kanneh, B.A., Tayleur, C. 2018. Nest ecology and conservation of Western Chimpanzees (Pan troglodytes verus) in Gola Rainforest National Park, Sierra Leone. Primate Conservation (32): 133-139. [http://www.primate-sg.org/primate-conservation-32/ link]
 
<span id="R00635"></span>BirdLife International. 2018. Eat chocolate, save a rainforest - the Gola Cocoa Project tells you how. Online: https://www.birdlife.org/worldwide/news/eat-chocolate-save-rainforest-gola-cocoa-project-tells-you-how


BirdLife International. 2018. Eat chocolate, save a rainforest - the Gola Cocoa Project tells you how. Online: https://www.birdlife.org/worldwide/news/eat-chocolate-save-rainforest-gola-cocoa-project-tells-you-how
<span id="R00636"></span>BirdLife International. 2019a. Important Bird Areas factsheet: Gola Forests. Downloaded from [http://datazone.birdlife.org/site/factsheet/gola-forests-iba-sierra-leone  datazone.birdlife.org ] on 05/07/2019.


BirdLife International. 2019a. Important Bird Areas factsheet: Gola Forests. Downloaded from [http://datazone.birdlife.org/site/factsheet/gola-forests-iba-sierra-leone  datazone.birdlife.org ] on 05/07/2019.
<span id="R00637"></span>BirdLife International. 2019b. Forests of Hope site - Gola Rainforest National Park, Sierra Leone. Online: [https://www.birdlife.org/worldwide/projects/forests-hope-site-gola-rainforest-national-park-sierra-leone  www.birdlife.org]


BirdLife International. 2019b. Forests of Hope site - Gola Rainforest National Park, Sierra Leone. Online: [https://www.birdlife.org/worldwide/projects/forests-hope-site-gola-rainforest-national-park-sierra-leone  www.birdlife.org]
<span id="R00638"></span>Brncic, T.M., Amarasekaran,B. & McKenna,A. 2010. Final Report of the Sierra Leone National Chimpanzee Census Project. Tacugama Chimpanzee Sanctuary, Freetown, Sierra Leone. [https://web.archive.org/web/20160308223613/http://tacugama.com/census.html link]


Brncic, T.M., Amarasekaran,B. & McKenna,A. 2010. Final Report of the Sierra Leone National Chimpanzee Census Project. Tacugama Chimpanzee Sanctuary, Freetown, Sierra Leone.
<span id="R00639"></span>Davies, Glyn & Schulte-Herbruggen, Bjorn & Kümpel, Noëlle & Mendelson, Samantha. (2008). Hunting and Trapping in Gola Forests, South-Eastern Sierra Leone: Bushmeat from Farm, Fallow and Forest. 10.1002/9780470692592.ch1.


Davies, Glyn & Schulte-Herbruggen, Bjorn & Kümpel, Noëlle & Mendelson, Samantha. (2008). Hunting and Trapping in Gola Forests, South-Eastern Sierra Leone: Bushmeat from Farm, Fallow and Forest. 10.1002/9780470692592.ch1.
<span id="R00640"></span>Gola Rainforest National Park. 2019a The Gola Rainforest Conservation Centre (GRCC). Online: https://www.golarainforest.org/conservation-center [https://web.archive.org/web/20220122143832/https://golarainforest.org/conservation-center link]


Gola Rainforest National Park. 2019a The Gola Rainforest Conservation Centre (GRCC). Online: https://www.golarainforest.org/conservation-center
<span id="R00641"></span>Gola Rainforest National Park. 2019b. Online: https://www.golarainforest.org/about [https://web.archive.org/web/20190722035626/https://www.golarainforest.org/about link]


Gola Rainforest National Park. 2019b. Online: https://www.golarainforest.org/about
<span id="R00642"></span>Klop, E., Lindsell, J.A. & Siaka, A. 2008. Biodiversity of Gola Forest, Sierra Leone. Unpublished Report. Royal Society for the Protection of Birds, Sandy, UK, and Conservation Society of Sierra Leone, Freetown, Sierra Leone. [https://web.archive.org/web/20260825215338/https://golarainforest.org/ link]


Klop, E., Lindsell, J.A. & Siaka, A. 2008. Biodiversity of Gola Forest, Sierra Leone. Unpublished Report. Royal Society for the Protection of Birds, Sandy, UK, and Conservation Society of Sierra Leone, Freetown, Sierra Leone.
<span id="R00643"></span>Lindsell, J. A., Klop, E., & Siaka, A. M. (2011). The impact of civil war on forest wildlife in West Africa: mammals in Gola Forest, Sierra Leone. ''Oryx'', 45(1), 69-77. [https://doi.org/10.1017/s0030605310000347 doi:10.1017/s0030605310000347]


Lindsell, J. A., Klop, E. and Siaka, A. 2011. The impact of civil war on forest wildlife in West Africa: mammals in Gola Forest, Sierra Leone. Oryx 45:69–77.
<span id="R00644"></span>RSPB. 2019. The Gola Rainforest: Sierra Leone's first rainforest national park. Online: https://www.rspb.org.uk/our-work/conservation/projects/gola-rainforest-sierra-leones-first-rainforest-national-park/


RSPB. 2019. The Gola Rainforest: Sierra Leone's first rainforest national park. Online: https://www.rspb.org.uk/our-work/conservation/projects/gola-rainforest-sierra-leones-first-rainforest-national-park/
<span id="R00645"></span>UNESCO. Gola Rainforest National Park. Online: https://whc.unesco.org/en/tentativelists/5747/


UNESCO. Gola Rainforest National Park. Online: https://whc.unesco.org/en/tentativelists/5747/
<span id="R02204"></span>Kiribou, R., Tehoda, P., Chukwu, O., Bempah, G., Kühl, H. S., Ferreira, J., …, & Heinicke, S. (2024). Exposure of African ape sites to climate change impacts. ''PLOS Climate'', 3(2), e0000345. [https://doi.org/10.1371/journal.pclm.0000345 doi:10.1371/journal.pclm.0000345]






'''Page created by: '''APES Atlas Team''' Date:''' 2019-09-02
'''Page created by: '''APES Atlas Team''' Date:''' 2019-09-02

Latest revision as of 17:53, 15 October 2026

West Africa > Sierra Leone > Gola Rainforest National Park

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Summary

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  • Western chimpanzees (Pan troglodytes verus) are present in Gola Rainforest National Park.
  • It has been estimated that 270 (CI: 159-468) individuals occur at the site.
  • The chimpanzee population trend is unknown.
  • This site has a total size of 710 km².
  • Key threats to chimpanzees are poaching and habitat destruction due to logging and shifting agriculture.
  • A REDD+ project is being developed and implemented at the site with the support of the Royal Society for the Protection of Birds, which aims to ensure funds for the continuation of conservation work in the park and improve the management of natural resources.


Site characteristics

The site consists of three forest blocks (Gola North, Central, and South) situated in southeast Sierra Leone, which are linked to Liberia’s Gola National Park, forming a transboundary peace park. The forests in Gola Rainforest National Park are a catchment for the Moro, Mahoi, Mano and Moa rivers, providing an important water source for nearby villages (Barca et al. 2018). The area forms part of the Upper Guinean forest ecosystem, a biodiversity hotspot with a large number of endemic species. Close to 1,000 plant species have been identified in Gola Rainforest National Park, half of which are endemic to the Upper Guinea forest ecosystem. In addition to the western chimpanzee (Pan troglodytes verus), other threatened species inhabit the area, including pygmy hippos (Choeropsis liberiensis), pangolins, red colobus (Piliocolobus badius), and Jentink’s duiker (Cephalophus jentinki). Gola Rainforest National Park is also classified as an Important Bird Area (BirdLife International 2019a) and has been included in the tentative list of World Heritage Sites.

Table 1. Basic site information for Gola Rainforest National Park

Species Pan troglodytes verus
Area 710 km²
Coordinates Lat: 7.60 , Lon: -10.92
Type of site Protected area (National Park)
Habitat types Subtropical/tropical moist lowland forest
Type of governance

Types of sites Governance types Habitat types

Ape status

The chimpanzee population in Gola Rainforest National Park is estimated to be 270 individuals, which is equivalent to almost 5% of Sierra Leone’s estimated population (Brncic et al. 2010). The population trend is unknown.

Table 2. Ape population estimates reported for Gola Rainforest National Park

Species Year Occurrence Encounter or vistation rate (nests/km; ind/day) Density estimate [ind./ km²] (95% CI) Abundance estimate (95% CI) Survey area Sampling method Analytical framework Source Comments A.P.E.S. database ID
Pan troglodytes verus 2015-2016 0.6 Gola Rainforest National Park Line transects Barca et al. 2018 96 chimpanzee nests were recorded SLE0012
Pan troglodytes verus 2009 0.76 0.25 (0.14-0.43) 270 (159-468) Gola forest reserves and potential corridors Line transects Brncic et al. 2010 Population estimate based on a re-analysis of data presented in Ganas 2009 SLE0002, SLE0003
SLE0004, SLE0014, SLE0015, SLE0017, SLE0018, SLE0021, SLE0024, SLE0025, SLE0026, SLE0029, SLE0032, SLE0036, SLE0037
Pan troglodytes verus 2009 0.27 (0.18-0.42) 305 (203-458) Gola forest reserves and potential corridors Line transects Ganas 2009 as cited in Brncic et al. 2010 Survey effort: 173 km SLE0002, SLE0003
SLE0004, SLE0014, SLE0015, SLE0017, SLE0018, SLE0021, SLE0024, SLE0025, SLE0026, SLE0029, SLE0032, SLE0036, SLE0037
Pan troglodytes verus 2006-2007 0.102 Gola West, East, North, and an extension of Gola North Line transects Klop et al. 2008 Survey effort: 245.28 km. No evidence of chimpanzees found in Gola West

Sampling methods Analytical frameworks

Threats

Approximately 28% of Gola Forest has been subject to logging (Klop et al. 2008). Large-scale logging operations in Gola started in the 1960s and continued until the late 1980s (Klop et al. 2008). Forestry Industries Corporation (FIC) and the Sierra Leone Timber Industry and Plantation Company (Sileti) were the two main companies responsible for the exploitation of timber in Gola (Klop et al. 2008). The resulting impacts from these logging operations include an extensive road network in Gola North, and a large clearing in Gola East. During the 1991-2001 civil war in Sierra Leone, the Revolutionary United Front set up camps near Gola forest; however, a survey that compared wildlife abundances before and after the war found no evidence of serious negative impacts from the war (Lindsell et al. 2011). Current threats in the area include small-scale logging, bushmeat hunting, and shifting agriculture.

Table 3. Threats to apes reported for Gola Rainforest National Park

Category Specific threats Threat level Description Year of threat
2 Agriculture & aquaculture 2.1 Annual & perennial non-timber crops Present (unknown severity) Shifting agriculture due to increasing population growth around the area and scarcity of fertile land (BirdLife International 2019b) Ongoing (2019)
4 Transportation & service corridors 4.1 Roads & railroads High (more than 70% of population affected) Extensive road network in Gola North as a result of large-scale logging activities (Klop et al. 2008), which facilitates access to the forest. Ongoing (2008)
5 Biological resource use 5.3 Logging & wood harvesting High (more than 70% of population affected) Large-scale logging operations between 1960s and 1980s (Klop et al. 2008), Small-scale logging by local communities (BirdLife International 2019a) 1960-1989

Threats

Conservation activities

A conservation program began in 1990 by the Forestry Division of the government of Sierra Leone and the Conservation Society for Sierra Leone (CSSL). Since then, the Royal Society for the Protection of Birds (RSPB) became a supporter of the conservation projects in Gola, and the site is part of RSPB’s Forests of Hope Programme. Conservation activities were interrupted by the civil war but resumed after the return of peace in 2002 (Lindsell et al. 2011).

A REDD+ project is currently under development at the site. The project will use funds obtained from the sale of carbon credits to improve the management of the park, fund long-term conservation work, develop sustainable farming practices, and the establishment of a research station in the park.

Gola Rainforest National Park also has its own conservation center, the Gola Rainforest Conservation Center (GRCC), which hosts visitors and researchers. The GRCC also offers capacity building and environmental education activities to the staff of the park, students, and local communities.

Table 4. Conservation activities reported for Gola Rainforest National Park

Category Specific activity Description Implementing organization(s) Year of activity
1 Development impact mitigation 1.4 Farm more intensively and effectively in selected areas and spare more natural land Develop sustainable farming practices (BirdLife International 2019b) Ongoing (2019)
1 Development impact mitigation 1.8 Use set-aside areas of natural habitat (e.g., mining, logging, agricultural areas) Bordering the site, cocoa trees are strategically planted alongside tree and plant species that increase soil fertility and attract birds (BirdLife International 2018) Ongoing (2018)
2 Counter-wildlife crime 2.3 Conduct regular anti-poaching patrols Forest guards patrol and monitor the forest (Gola Rainforest National Park 2019a) Ongoing (2019)
4 Education & awareness 4.1 Educate local communities about apes and sustainable use The Gola Rainforest Conservation Center offers environmental education and conservation awareness to school classes and local communities (Gola Rainforest National Park 2019a) Ongoing (2019)
5 Protection & restoration 5.2 Legally protect ape habitat The site was designated as National Park in 2010 (Gola Rainforest National Park 2019b) 2010-Ongoing (2026)
7 Economic & other incentives 7.2 Provide non-monetary benefits to local communities for sustainably managing their forest and its wildlife (e.g., better education, infrastructure development) RSPB is developing a REDD+ project which aims to provide funds for the long-term management of the site and support local people in the sustainable management of resources (RSPB 2019) Ongoing (2019)

Conservation activities

Challenges

Table 5. Challenges reported for Gola Rainforest National Park

Challenges Specific challenges Source Year(s)
2 Resources and capacity 2.5 Lack of equipment/transportation Brncic et al. 2010

Challenges

Enablers

Table 6. Enablers reported for Gola Rainforest National Park

Enablers Specific enablers Source Year(s)
1 Site management
2 Resources and capacity
3 Engaged community
4 Institutional support
5 Ecological context
6 Safety and stability

Enablers

Research activities

Several surveys have been done at the site to monitor wildlife and estimate species abundances (e.g., Brncic et al. 2010, Klop et al. 2008, Davies et al. 2008), and the nest ecology of chimpanzees has also been studied at the site (Barca et al. 2018).


Documented behaviours

Table 7. Behaviours documented for Gola Rainforest National Park

Category Behavior Observation type Prevalence Observation dates Source Habituation Species Comment
Not reported

Behaviours

Exposure to climate change impacts

As part of a study on the exposure of African great ape sites to climate change impacts, Kiribou et al. (2024) extracted climate data and data on projected extreme climate impact events for the site. Climatological characteristics were derived from observation-based climate data provided by the Inter-Sectoral Impact Model Intercomparison Project (ISIMIP, www.isimip.org). Parameters were calculated as the average across each 30-year period. For 1981-2010, the EWEMBI dataset from ISIMIP2a was used. For the two future periods (2021-2050 and 2071-2099) ISIMIP2b climate data based on four CMIP5 global climate models were used. For future projections, two Representative Concentration Pathways (RCP) were used. RCP 2.6 is a scenario with strong mitigation measures in which global temperatures would likely rise below 2C. RCP 6.0 is a scenario with medium emissions in which global temperatures would likely rise up to 3C by 2100. For the number of days with heavy precipitation events, the 98th percentile of all precipitation days (>1mm/d) was calculated for the 1979-2013 reference period as a threshold for a heavy precipitation event. Then, for each year, the number of days above that threshold was derived. The figures on temperature and precipitation anomaly show the deviation from the mean temperature and mean precipitation for the 1979-2013 reference period.

The estimated exposure to future extreme climate impact events (crop failure, drought, river flood, wildfire, tropical cyclone, and heatwave) is based on a published dataset by Lange et al. 2020 derived from ISIMIP2b data. The same global climate models and RCPs as described above were used. Within each 30-year period, the number of years with an extreme event and the average proportion of the site affected were calculated (Kiribou et al. 2024).

Table 8. Estimated past and projected climatological characteristics in Gola Rainforest National Park

Value 1981-2010 2021-2050, RCP 2.6 2021-2050, RCP 6.0 2071-2099, RCP 2.6 2071-2099, RCP 6.0
Mean temperature [C] 26 27.1 27.1 27.2 28.3
Annual precipitation [mm] 2286 2245 2316 2290 2324
Max no. consecutive dry days (per year) 38.7 35.5 30.9 30.5 35.5
No. days with heavy precipitation (per year) 6.3 6.8 6.7 6.9 8.5


Table 9. Projected exposure of apes to extreme climate impact events in Gola Rainforest National Park

Type No. of years with event (2021-2050, RCP 2.6) % of site exposed (2021-2050, RCP 2.6) No. of years with event (2021-2050, RCP 6.0) % of site exposed (2021-2050, RCP 6.0) No. of years with event (2070-2099, RCP 2.6) % of site exposed (2070-2099, RCP 2.6) No. of years with event (2070-2099, RCP 6.0) % of site exposed (2070-2099, RCP 6.0)
Crop failure 7 0.12 8 0.08 7 0.09 14 0.08
Drought 0 0 0 0 0.5 12.5 0.5 12.5
Heatwave 9.5 67.71 10.5 77.78 10.5 83.62 19 86.31
River flood 1 2.09 1.5 2 2 2.19 5.5 3.18
Tropical cyclone 0 0 0 0 0.5 0.14 0.5 3.57
Wildfire 30 0.5 30 0.48 29 0.5 29 0.49


  • Precipitation anomaly in Gola Rainforest National Park
  • Temperature anomaly in Gola Rainforest National Park

External links

References

Barca, B., Turay, B.S., Kanneh, B.A., Tayleur, C. 2018. Nest ecology and conservation of Western Chimpanzees (Pan troglodytes verus) in Gola Rainforest National Park, Sierra Leone. Primate Conservation (32): 133-139. link

BirdLife International. 2018. Eat chocolate, save a rainforest - the Gola Cocoa Project tells you how. Online: https://www.birdlife.org/worldwide/news/eat-chocolate-save-rainforest-gola-cocoa-project-tells-you-how

BirdLife International. 2019a. Important Bird Areas factsheet: Gola Forests. Downloaded from datazone.birdlife.org on 05/07/2019.

BirdLife International. 2019b. Forests of Hope site - Gola Rainforest National Park, Sierra Leone. Online: www.birdlife.org

Brncic, T.M., Amarasekaran,B. & McKenna,A. 2010. Final Report of the Sierra Leone National Chimpanzee Census Project. Tacugama Chimpanzee Sanctuary, Freetown, Sierra Leone. link

Davies, Glyn & Schulte-Herbruggen, Bjorn & Kümpel, Noëlle & Mendelson, Samantha. (2008). Hunting and Trapping in Gola Forests, South-Eastern Sierra Leone: Bushmeat from Farm, Fallow and Forest. 10.1002/9780470692592.ch1.

Gola Rainforest National Park. 2019a The Gola Rainforest Conservation Centre (GRCC). Online: https://www.golarainforest.org/conservation-center link

Gola Rainforest National Park. 2019b. Online: https://www.golarainforest.org/about link

Klop, E., Lindsell, J.A. & Siaka, A. 2008. Biodiversity of Gola Forest, Sierra Leone. Unpublished Report. Royal Society for the Protection of Birds, Sandy, UK, and Conservation Society of Sierra Leone, Freetown, Sierra Leone. link

Lindsell, J. A., Klop, E., & Siaka, A. M. (2011). The impact of civil war on forest wildlife in West Africa: mammals in Gola Forest, Sierra Leone. Oryx, 45(1), 69-77. doi:10.1017/s0030605310000347

RSPB. 2019. The Gola Rainforest: Sierra Leone's first rainforest national park. Online: https://www.rspb.org.uk/our-work/conservation/projects/gola-rainforest-sierra-leones-first-rainforest-national-park/

UNESCO. Gola Rainforest National Park. Online: https://whc.unesco.org/en/tentativelists/5747/

Kiribou, R., Tehoda, P., Chukwu, O., Bempah, G., Kühl, H. S., Ferreira, J., …, & Heinicke, S. (2024). Exposure of African ape sites to climate change impacts. PLOS Climate, 3(2), e0000345. doi:10.1371/journal.pclm.0000345


Page created by: APES Atlas Team Date: 2019-09-02